the more i hear about this, the more i am inclined to go with the genetics, when human parents give birth to a deformed baby , they often dont exhibit any signs of the deformity that their child did. on the other hand, a now known d3 deficiency is a huge flag ,there could be something that is inhibiting d3 utilization even though there is plenty given. do these animals also get sun? if so , it is possible that somehow the exposure to natural sunlight is somehow conflicting with your regimen in a way as to prevent d3 utilization (may be an excess of vitA)also are the animals fed strictly from known wholsome feeders or do they also have access to outside feeders that may have had some pesticide exposure ? i realize that all of this speculation only increases the frustration level, and that you need to get a sense of what went wrong if you are ever to consider breeding this animal (probably not, given its suspected liver problem?) or even any of its appearent healthy (non cleft) offspring, do you also have another female that is gravid ?and if so, did your animals come from the same source? if it came from the same source and the same thing happened again , it could still be read two different ways, (although i would tend to lean towards the genetics), but if your other animal came from a different source , it would tend to indicate more of a regimental thing. (unless they both used the same sire, which would confuse things all over again ) i was wondering was it your goal to breed jax, or was this just something that happened ? the sad reality is, you may never be able to pinpoint the exact cause and IF it is your goal to breed jax then you might be better off starting from scratch, meaning new animals from different source and a complete rethink of your entire regimen (including light) from the ground up, i know this has got to be hurtfull as well as frustrating ,so i hope you are eventually able to reach some sort of conclusion, and what ever that conclusion is , i sincerely hope it is able to offer you some closure on the matter. even the best keepers sometimes have problems that dont make any sense, or are unable to figure out , all any of us can do, is do our best , try to act responsibly (both of which you have clearly done), and hope for the best. this is a good example to all keepers, that none of us know as much about these animals, as we think we do. just when you think everything is under control is when the most difficult of problems seam to arise