As in, all knobbed out, getting nobbly, or, Look at those knobbules!!!
The ones on the coronal process(their heads) are part of the skull itself, as with other chameleons. (I know, Ive taken them apart)
The ones on their appendages, are just how the scales are. Its their particular scalation.
Off the top of my head, I would assume that it is a trait that has evolved to maximize surface area of the scale, while keeping the overall package small. (nothing wrong with that, its the motion of the ocean that counts...right?)
Anyhow, this would allow the animal to absorb more sun, while keeping the water and food needs to a minimum.
These are small chameleons, that live in brushery and shrubbery and whatnot. So to me, that seems to make sense.
I could be completely wrong though, as its just an assumation, and I am in no way qualified to make a direct official statement on the matter.
Seems like a good question actually. Lets ask Chris! I bet hell shoot us straight...