“On inclines and declines, limb placement was more variable than on level substrates. The tail never contacted the substrate during level locomotion; however, on inclines and declines, the tail was held closer to the substrate, with short substrate contacts in one-third of the analyzed trials. Regardless of substrate orientation the tail was always held straight above the branch; therefore, rotational moments induced by the tail were minimized.”…
https://journals.biologists.com/jeb.../Biodynamics-of-climbing-effects-of-substrate
https://journals.biologists.com/jeb.../Biodynamics-of-climbing-effects-of-substrate
