“The F. oustaleti and F. verrucosus complexes are well supported as sister clades with mitochondrial data, and as distinct nuclear clusters with both nuclear genes analyzed. While there is substantial genetic variation recovered with mitochondrial data, there is little variation within each species complex with the nuclear genes”…
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4892496/
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4892496/